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You can switch back and forth whenever you like once they are unlocked (even mid fight), but you have to go into the menu each time which is pain.
Genranlly it better to buy the one upgrade on each main skill first and then go back for second and thirds after having one one each.
Either you get arrows, or you can brand, but not both at the same time.
I guess I am not far enough along for runes yet, but I will keep that in mind when I get that far. I do like the full health with a drain so if I can get regular arrows some other way that will be great.
You mean a piece of gear :) The rune system from shadow of mordor is no more.
OH. I've been considering those as being different rings, my bad :P