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No idea if it has gameplay impact.
Yes, totally, but please don't. That's just messing with my stress level. Just... don't.
Other than being a skin feature adding some additional incentive to unlock the skull sets it doesn't do anything else as far as I know. Beating the game with crystal skulls on medium or hard doesn't give another ending, for example.
The skulls kind of fit the story thematically (the endless loop the crew find themselves in).