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Jetfish only eat fruits and other plants(they also eat leeches). Squidcadas only batflies, leeches and centipedes.
Pole plants and kelp can also eat bats. Kelp also eats fruit and other things you throw at it when it charges at you.
There's also a DLC lizard that eats bats I think.
Small spiders also eat bats.
Lizards will attack bats, but you see this fairly rarely because they're usually not within range. That being said, I've seen a magenta lizard grab and run off with a dead batfly that I had hit with some rubble a few moments before.
Aside from that and other things already mentioned, technically anything edible to survivor is also edible to slugpups, though those are technically still slugcats in the end.