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Right where it belongs.
From hardest to easiest per floor:
1) Gatling Gull > Trigger Twins > Bullet King
None of them are really threatening, but i find that getting trapped in close quarter with the boss is the main reason for taking damage against them, and Gatling Gull seems to excel at this.
2) Ammoconda > Beholster > Gorgun
Beholster and Gorgun are both predictable and straightforward (especially from a bullet hell aspect), giving you a lot of space and time to react. Gorgun is a bit annoying with the screamcircles, but Beholster is more threatening with his minions and rockets though you can still deal with them by having them run into a wall and rolling the opposite way. Ammoconda on the other hand... fast bullet spam, very erratic, and its "allies" make the fight much harder. But i think the biggest offender is how it sometimes charges at you way too aggressively, its massive body reducing a lot of your room for dodging.
3) Treadnaught > Cannonbalrog / Mine Flayer
I'm unsure which one i'd rank higher between those last two. I find them both easy but capable of cheap shots. I've always had cover on the Mine Flayer room though, which makes pretty much the whole fight cheesable. The bell attack seemed pretty easy if you take cover, or go at the bottom of the room instead of the sides. From the side, the attack is harder to tackle, but still feasible. All the other attacks aren't overly fast, give you space, and the minions give you all the time in the world to reposition yourself.
Cannonbalrog can corner you, or surprise you with the "darkness" attack, but aside from that... it's all easy patterns to dodge.
Treadnaught is absolute chaos. There's a massive ammount of random bullets everywhere, you are forced to constantly move around a very clunky room design, the minions can hit you in the back with a cheap shot or add to the bullets, and the fast exploding bullet is my bane. It comes at you, fast, with a huge explosion radius, while you're trying to navigate the room to not get trapped or overwhelmed at the worst times. It's always this or a stray bullet.
4) Hard to make a choice... I feel like they are all pretty balanced and equally challenging, depending on your playstyle. Kill pillars is probably on the easier side though once you get a feel for their attacks, while Wallmonger on the harder side.
Havn't fought secret bosses and pasts in a while so i'm not too sure anymore, but as long as you know what to expect in a past and how to handle it, none of them are too bad. Bullet past surprised me at first, i lost most of my health really quick, unfamiliar with the boss... but once i figured that i could just take my time, slowly sidestep the bats, and just roll in after some patience, smack the boss once or twice, roll out, it pretty much secured my first attempt. I found the robot harder. Can't recall the others, but i think Marine/Pilot gave me the most trouble, followed by Convict, Robot and Huntress. Don't remember if i failed any past, but probably 1-3 of them, once or twice.
Chamber 2: Ammoconda
Chamber 3: Mine Flayer
Chamber 4: Wallmonger
Chamber One - Old King, Trigger Twins, Gatling Gull.
Chamber Two - Gorgun, Ammoconda, Beholster.
Chamber Three - Treadnaught, Mine Flayer, Cannonbalrog.
Chamber Four - High Priest, Wallmonger, Kill Pillars.
-Gatling Gull is just tough to outrange in floor one most of the time, and if you get the room with the bushes, you better start praying.
-Beholster, at least for me, is rough. None of my guns ever seem to do enough to shoot down the homing rockets in one shot, and while focusing on those, I either get hit by his minions or other projectiles.
-Cannonbalrog is the only one of these bosses to actually claim a life on me so far, just because of his annoying invisible darkness tactics.
-Kill Pillars, I have only ever beaten once, maybe twice. If you get past the inital circle of death, it isn't horrible, but the Jammed bullets took me a while to understand the pattern. Once you kill at least one, it gets much easier.
I haven't gotten to beat the pasts of the Convict, soldier, or robot yet, but between Pilot and Huntress, Pilot is harder by a mile. Huntress was a cake walk.
Different strokes, I guess.
My choices are Bullet King 1, Ammoconda 2, Mind Flayer 3, and the Marine's Past. Granted I'm still 3/6 on the Pasts, but the Marine's gave me the most trouble. The Hunter and Pilot needed two a piece - one to learn what to do and the other to win - while the Marine took me like six :( My answer might change to the Bullet's Past later, 'cause the one time I reached the Past with the Bullet I was curb-stomped by Cannon lel
2. Ammoconda. His attacks are among the most unpredictable in the game.
3. Cannonbalrog. I don't get all the hate for Mine Flayer, most of his attacks have huge safe spots? But cannonbalrog has the attack where he just spits out tons of bullets in all directions with no preplanned safe spots, and the attack where he spits a rotating stream from each eye that you have to dodge roll, and he can get up in your face before either.
4. Kill Pillars. Many of the attacks fill the room with tons of bullets in not-very-reliable patterns thanks to coming from 4 places at once, and the attack where they chase you and try to stomp on you just feels unfair.
Past: Pilot's past. Maybe I'm just not as good at that kinda game, but its attacks blow the other pasts out of the water.
The Gull is overall a hard boss and the trigger twins are prety random so at times the fight can be a pain due bad rng.
Beholster / Ammoconda
basically the same, Beholster has hard pattersn while ammoconda is rng hell
Mine Flayer/Treadnaught
Same thing again, Mine flayer is not that hard except for that damn bouncing bullet thing, treadnaught in the other hand is again rng hell, sometimes is a joke, sometimes is pain
High Priest
Not a lot to say here, he is just hard.
2. Ammoconda because you know why. Beholsters trivial as long as you control his adds and missiles, and gorgun is a joke.
3. Cannon Balrog when he starts the fight with an attack that moves toward you and you start to at the left or right side of the arena. If this happens he essentially forces you to blank or get hit to reposition so that he can't pin you against the wall. The mineflayer is largely pattern based though his patterns are pretty intimidating and demand you to play well and understand the fight. Treadnaught is pretty simple as long as you clear the adds and maintain a good amount of distance from him.
4. The high priest. His Personal attacks on the surface aren't terribly hard to deal with but he has these random bullets that shoot from the side of the arena that can screw you over. The wallmonger is very good at punishing players who don't know how to fight him but once you learn his attacks he's a huge push over. The kill pillars are only really threatening when they try to chase you down but even then if you just focus on weaving you can clear them very easily. Also they're super susceptible to aoe attacks.