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Edit: I think all the predator fish take up 4 slots. I transfered a Crab Squid and a Bone Shark from one double stack containment to another stack the same size to keep my 2 Cuddlefish company. :)
That was what I suspected. Each predator takes up 3x3 (9 slots!), and my 30-population limit tank is probably overfull by now. Fortunately eggs still hatch, and excess fish don't die. They just don't reproduce.