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He said it can happen when he first places the pens or when he logs off.
He's verified there is water and appropriate food (capybara feed, turkey feed, and meat for the angry fish).
He has a breeding pen and a tribesman assigned to it. He changes who is assigned to the breeding pen every time it happens. It still happens.
He has built new pens 3 times and it still happens.
He has changed where they're placed and it still happens.
I found this on Reddit and showed it to our server admins: https://www.reddit.com/r/PlaySoulMask/comments/1dm4ld1/animals_in_pens_disappearing_on_dedicated_server/
They made the change and he is going to try again today. Because I personally toured his base and I can tell you, his base is fine, the base is not the problem. Everything he has is covered by his fire. And the area is also fine as my husband & I ironically have our own base just next to his spot on our personal server and there are no issues.
Will update you if that works.
Keep one of each fish in the Llama basket overnight, then try again.