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(I'm guessing you meant victory lap and not rerun - but I thought I should clarify).
Also there's no point in getting keys for mega satan until the lap where you are the lost as they will be used in the lap they get picked up on - as for dad's key and the like they are likely to vanish on the 3rd lap.
Ye, actually meant victory lap not rerun, my mistake here, sorry
Did u beat the Lamb in your victory lap as other character? (lets say u kill him first time with isaac and in victory lap u died and got revieved as blue baby by ankh and got achievement 'kill the Lamb as blue baby' after u killed him in victory lap)
No matter what if you die and are revived as ??? with Ankh you then get ??? unlocks - I assume you stay as ???, or any other character you were, on the other laps before you turn into the Lost.
So if you get to lap 2 and revive as ??? you will get his hard boss marks - then turn to The Lost on lap 3.
If you turn into ??? on lap 1 and beat the lamb you will get that boss mark and lap 2 should start you as ??? I believe - but I've never tried that myself.