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Other than that it mostly comes down to the proximity of human habitation or the presence of bioluminiscent flora like Mindshrooms. So even the more isolated and low traffic areas of lower underrail will have some ambient light from all the train tracks, stations and settlements while the lower caves or completely isolated areas like the Black Crawler's habitat are completely in the dark barring the presence of shrooms or camp fires.
The Black Sea is somewhat unique because it's a wide open area with a dizzyingly high cealing (by Underrail standards) and there's light coming from the pirate bases, the expedition camp, native villages, the mutie settlement and even the glowing fang. Don't know how strong these light sources would realistically be but they at least keep the Sea from being pitch black unless, again, you crawl into some hole or cave.
I think the game does a decent job of taking these factors into account and having diverse lighting levels but it can't be completley realistic or it'd be a pain to play.
I'd hate to have to light flares all the time, man. Stealth characters'd pretty much need night-vision if the darkness were realistic.
It'd really suck if the whole game looked like you couldn't see 5 meters in front of you, but I think the game could use more ways of conveying the relative darkness, compared to what people in Underrail are used to.
I have to assume that after so many generations of living underground, people are generally more cave-adapted than we are, but most tunnels seem dark enough to penalize ranged attacks at least
Bioluminiscent fungi hidden in the invisible ceiling, of course!
In all seriousness it's never explained, but the above could be true, if Mindshrooms are anything to go by. Like others have said it's just a gameplay thing.