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As an added note... definition:
humanity
noun hu·man·i·ty \hyü-ˈma-nə-tē, yü-\
: the quality or state of being human
: the quality or state of being kind to other people or to animals
: all people
There is a threat to a PEOPLE(Culture AKA Hiigar.ians? and the Kudash.) though, and Humanity is a refrence to a people.
Just... watch the cutscenes
Pretty sure humans are the biggest threat to humanity, regardless of an alien threat.
While the trailer does look like it was made by a spastic thirteen year old hopped up on Mt. Dew, it does not hold any grand meaning to the game itself.
It is implied in Homeworld 2 that all of the races involved in the story descend from the Progenitors, a long extinct race that seeded the galaxy with life and whose hulking derelicts litter some regions of space. This is also confirmed in the development documents of the original Relic team.
It's obvious Gearbox just wanted to go "Michael Bay" with the trailer, but personally I think all they managed is to confuse people who're not into it only for some nice explosions.
Could also very well be that the galaxy the story takes place in isn't even our galaxy. The progenitors "arrived" in this galaxy a long time beforehand. Intergalactic travel is certainly possible - the Bentusi can do it in dire circumstances.
And yes, no aliens to my knowledge. some of them look pretty naffy, like the Bentusi, but they are still recognizeably (trans)human at a basic level. Here's a Bentusi http://img1.wikia.nocookie.net/__cb20060512165817/homeworld/images/8/84/Bentusi.jpg
Remember you're dealing with hundreds of thousands, possibly millions of years of divergent evolution (and deliberate modification in the case of the Bentusi) so yeah, things change. Any good sci fi writer incorporating deep-time will tell you that the one thing you must keep consistent is the eyes, as that is the mirror of humanity. Homeworld does this well.