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If there is a green tick, it means that the said country will join the war as an ally. If it shows a red cross, they will refuse the call to arms.
If you make an ally co-belligerent, it makes them a direct target of the war, rather than just an ally. That way, you can take land from them with less penalties. However, it also means that co-belligerent countries can also ask their own allies for help.
The attacker decides who is belligerent because it's the one who declares the war. The defender has no say in who is the primary target of the war. It's like while getting mugged on the street, you say "I'm not the one you are mugging, it's my friend over there." HRE members are the only exception.