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Also, yes. The story is unlocked by progressing through the Solar System. Unlocking the Operator is part of that, however, unlocking the Operator's set of abilities demands playing in the Plains of Eidolon, Zariman/Circuit, Sanctuary Onslaught or the use of Lenses to obtain Focus and upgrade the Focus Schools.
Likewise, appearance customisation is very limited without unlocking additional outfits. Most of these can be obtained in the open-world syndicates and crafted from materials, but then quite a lot of cosmetics are accessed only through the in-game Market for Platinum.