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You have the Attributes like STR,INT etc which increase inflicted Damage,increase Healthpoints etc.
Below are the Skills (its more like Skillsets) that enable you to use "Combat Arts" (those are the things with those 3 bubbles below each icon.So,if you put points into the "Lost Fusion" Skill,it will lower the cooldown of all combat arts that are under the Lost Fusion tab and raise the maximum level of combat arts you can use without getting a malus.
You boost each Combat Art individually with those Runes you find.Lets say you find a chest full of runes for "Jolting Touch",boost it up to lvl 20 while your character is only lvl 2.
You´d get a cooldown that is unbearable on that exact combat art.
As said above,to avoid that you have to put points into the SKILL that handles the CombatArts set where "Jolting Touch" is located.
You do not get more usable Combat Arts,there are 3 sets for each character class with 5 usable attacks/buffs.They become usable as soon as you use the first rune with that Combat Art name if i recall correctly.
The quickbutton for Healthpoitions is the Spacebar,picking up loot around you is on Q but i believe you can change that in the control options.
Hope that makes at least a bit of sense/is readable & helpful
Most of those runes are useless because the dont belong to your class (you cant read them anyway but you can forge them into equipment slots) - but dont sell them. Instead go to a Rune Master NPC and exchange them for one of the runes you need (you can choose there which one you want).
Exchanging 1 to 1 is very expensive but 4 useless runes for one rune of your class very cheap, so you just gather a lot of useless runes and then go to a rune master.
Be careful with leveling up combat arts because the regeneration times increase.
http://www.sacredwiki.org/index.php/Sacred_2:Runes
There comes the skills into play. If you level up you gain points you can put into skills,
and choose new skills. You need to put the points you get when you level up into the skill that lowers the regenerations time of the combat arts you use most.
Ever character has three combat art groups and every combat art group has one skill that lowers the regen times.
http://www.sacredwiki.org/index.php/Sacred_2:Skills
For example if you have a "dragon mage" and you use a combat art of the "elemental magic" group often you need to choose the skill "Elemental Magic Focus" at soon as possible and regularly put points into it, so that you can level up the combat art without problem.