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do you use a modifier?
I think that this is not related to the missions previously created ..
I have had that issue, sometimes the profile will lose the added modifier button/toggle from the HOTAS, and any command that you programmed with that modifier will show a red exclamation mark. I use about 4 modifiers on my HOTAS, so I had red exclmation marks everywhere until I re-added those missing modifier buttons/toggles from the HOTAS.
So look at the ones with red exclamation marks, see what the programmed HOTAS buttons are or whatever, and see if you need to re-add it as a modifier.
Just a guess.