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Here's the lore based answer:
No, space marines keep their helmets on during combat.
This game is however based on the tabletop wargame Warhammer 40'000, where you fight out big battles with your hand-painted minis (basically plastic toy soldiers). The whole helmetless space marine thing came from the fact that each box of space marines came with a few options of helmetless heads you could use for your officers and unit leaders. When you have a hundred tiny figures on tne table, it is much easier to determine at a glance which of them are your heros and special characters.
The whole thing got so prevalent, that in the rulebooks space marine heroes often were depicted without helmets for dramatic effect, since a square-jawed hero yelling in the middle of the firefight looks much more badass than just another guy with the same helmet as all the others around him.
However in the novels, even the heroes wear their helmets like the rank-and-file space marines, and they only take it off when they have to interact with normal humans in a non-combat environment.
Nowadays, the newer model kits often feature extra helmet bits specifically designed to be glued to the model's belt, so you can have the option of your hero going bare-headed while also indicating that he in fact has a helmet, and he didn't just forgot it on the space ship before the mission.
As far as I know, in the campaign mode, there is a setting where you can choose to have the characters either:
- always have their helmets on
- always have their helmets off
- only have their helmets off during cut-scenes
From a bodybuilding perspective it's not plausible they would have low T with the size of them. They almost certainly have some insane T level that would make the worst steroid abusers look like nothing lol.