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I think you have to finish an entire stage (10 floors boss) with the weapon at that level. Im not sure if it has to be actually held into the boss fight but that is what I did do make sure they saved.
You can replay and grind earlier levels at any difficulty to get what you need and use the 'refresh inventory" to make it go faster.
This is exactly what I tried to say. I did this with the smg, but the game didn't save the level of the weapon. I can still only find a level 2 smg in the shops.
The shotgun I did beat a boss with and the upgrade did not stay. The Railgun I just got to stay at level 3 without beating a boss, same for Fire Katana.
I think this might be it if I'm remembering correctly. I do believe the time I got the Shotgun and Railgun to level 3 without having them save I got both level 2 and 3 together on the same run.
This might indeed be the culprit of the issue. I got the smg to stay at level 3 now after finishing another boss with it. I will test more with different weapons soon since, well, I want to have all of them at level 3 of course.