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They are a one time thing per difficulty (easy, normal, hard). The rewards at least, You can do it again if you missed some of the hidden boxes/chests inside
It's also very very challenging once you start doing the Hard Mode be warned.
It basically a Solo Challenge and the CS Requirements is a suggested Warning though some can be achieved whilst below the CS Requirements.
I can't imagine what normal and hard are going to be like.
While getting higher level you will unlock new content and things to do. Both solo and co-op. Every monday there is weekly activities resets for replay-ability.
The main source of relic shards on:
1) Crew shop
2) Chance on 7 x Claire's dream every week
3) Bought in Build mode that reset every week
I believe you can still farm them, even without chips.