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Okay, so first you have to equip them before you start a run, then you use them instead of picking a new ability when you level up. it seems there is a very limited number of summons per run too, so don't use them right away.
EDIT - Erroneously stated you couldn't re-summon the same demon, but looks like you get one summon per "slot" you've unlocked and can use them all on the same demon if you want (though that means you aren't leveling the unused ones during that run).