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You unlock them in the altar of hope, 8 candles per unlock.
Yes, it's a game over. It's like "This run sucks! Let's restart!"
Your permanent upgrades are hero shrines (Unlocking abilities.) and candles are used to earn items you can find in the world, heroes, etc.
But the chapters don't connect to each other, do they?
Each chapter would be like a different dungeon, while the shrine would be like the hamlet.
The main difference being that you don't have a permanent roster of heroes and equipment like in the first.