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If the fish is clean, healthy, fed and happy, and it's not night, but *still* has the warning light lit up, that is when you hit the Discipline button. That'll raise its training meter and turn off the warning light.
That's... about all I know about the Gyao system. I had a Dolphin as my first, and i kept training it wondering if it'd grow further, but it eventually died (old age? neglect? i'm not entirely sure) and my second grew into a Beluga. I'm unsure if it's optimal to grow them up then let them die, but that's what it feels like to me. Which seems a little odd.
Yea I assume if your just trying to unlock them all it's best to just treat them well until they evolve into their first form then let them die, but apparently they can evolve a 2nd time someone said in another thread, but I treated my first one (Dolphin) perfectly and it just up and died of old age I suppose. Though I was never able to get it's training very high as it almost never had a Warning to Discipline without a reason.
https://steamcommunity.com/id/laij4/screenshot/2003587608399579464/