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Once you kill the boss in the region, next time you load into that region, a nymph will spawn somewhere on it.
The post-boss nymph spawn locations from memory:
Upturned Forest - The far corner of the map, there's a bit you have to drop down to get to, one of the rooms in there (hard to explain)
The Fault - The room full of shipping containers.
Great Forest - At the end of the initial area, after the broken drone with a spotlight pointed at it the open rooftop area where you find the bazooka.
Arboreal Nest - A random room relatively early on in the "main route"
Accumulation Zone - The big empty plateau area, past the earliest laser shotgun worm things you can encounter, i think it's where you find the double barrel shotgun?
Forbidden Domain - In the boss room.
Also there have been a few nymphs in the areas that were locked behind the red forcefield barriers
The Depths: Theres one towards the start at a left bend at the first fork in the road
Tomb of Divine Wisdom: Left of the black water pool
(PS anyone know where the tank lady goes after the tomb?)
Also found a few more:
Central Cavern: in the room through the red barrier next to where you first meet Yoyo (the merchant at your base)
Water Treatment Levels: in the middle of the ramp between the resevoir area and the tower the boss is in
The Bridge: go through door 06, up the stairs on the right, up the elevator, and out into the area with the shipping containers
location guide here
https://www.youtube.com/watch?v=z3MICpk9mFk
First tree you met in this game, she sits nearby
Funny huh? Same ♥♥♥♥ with Grand blue fantasy relink, you grind the fk out of that game only to get the weapon at the end when you don't need it anymore. I had this same problem with this game, but at the 2nd stage of the upper level node you buy a flying frame when you've basically already completed the level.