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It is revealed later that Circe has the ability (granted to her from the 2nd capture by the Jewel Thief in Gaiden) to turn the male soldiers into C.C. that obey her commands.
It is done through intercourse with the male soldiers. So when they became C.C. their bodies regenerate from death. Essentially means Circe had intercourse with everyone who turned into a C.C. and bleeds blue blood throughout the past 9 years since the Cathedral had been built. Though they still emulated the behavior of their original host until the brain is shot. They regenerate, but anything that was part of the human original personality is killed. That's why Callaghan and the other soldiers act as puppets at the end. They regenerated and are now only C.C. that can follow orders without any of their previous human personality.