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The skill tree upgrades are almost exclusively not just raw stat upgrades. Stuff like dash retaining perfect launcher, teching, bonus rarity on perfect clear, healing hp when you enter every room, etc. The only real raw stat upgrades are +50 starting hp and +100 starting dust.
Mid run the upgrades also have pretty significant changes. Slows, dots, super armor, speed up, back attack boost and crit chance. All the bolts play different as well. You might even change up your gameplan if you get an on hit style blessing (like combustion or lightning shockwave). Then theres synergy boons as well.
Judging by your wording, I'm guessing you just aren't using aerials enough to extend your combos.