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Once you spend the mana to unlock the spell you can use it right then and there. You have to do the challenges for each spell to increase their damage and increase that spell color's power.
Yes, some spells require you to go to certain places and do certain objectives to obtain them.
No, the entire game doesn't have just 1 tree .. there's 4 tree's that you unlock as you play through the story.
Yes, you get better modes of transportation. You can cover vast amounts of area very quickly once you get them.
When you get mana you can use it to learn a skill. This mana can be, indefinitely, refunded to use on another build. Think of it kind of like Code Vein or something where you can swap builds whenever. You can do challenges to gain permanent stat buffs and bonus effects. The challenge unlock is acquired via the book in archive where you can then access at any rest location that isn't a field bonfire. Completed challenges are permanetly completed even if you unlearn and later relearn a skill.
One skill with parkour allows you to dash up a wall to go even higher with an extra jump dash acquired extremely early on in basically tutorial. Once you get used to the stamina system and unlock an additional 4th stamina bar it becomes much more manageable, and of course knowing how to properly parkour (like how to properly drop to lower areas/ledges scaling down without losing much speed or bursting stamina bar) or tricks like using the first attack while falling can negate fall dmg entirely if done just before landing making zero stamina drain on landing from any height. There are tricks like a parkour attack can instantly charge your R2, keep your sub skill set to your desired skill to spam (it charges faster with combat actions, briefly swap to others and use them then go back to preferred sub skill), etc.
I will warn you that to unlock the next tree after Frey's starting one you would want to rush to it and then do it then go back to explore if this is something you want ASAP because the area to explore is actually friggin massive (its not quite Just Cause 3 or whatever type large, "maybe?", but its pretty huge). This can heavily pad the time between the next skill tree unlock if you do that stuff first so something to consider if this is a core goal. That said you can become a murdering machine with just the first tree alone ripping apart even mutants like the tanky Goliath after challenges unlocked, gear upgraded, etc. to the point they can't really properly even fight back (esp with the knock down bonus on the pre-order DLC on top of it all).
The environment design is MUCH more interesting, at least imo, after the initial grass plains. Basically, tutorial > city cipal > minor story event to get someone > city > then unlocks area south of city cipal which after a brief travel through this one region leads you to the desert region. Much better from there (in fact, even compared to the initial desert area it gets yet better in subsequent desert zones and such).
Don't stress yourself about getting everything discovered if you don't want to asap. Some stuff can't even be accessed such as high up stuff due to needing a mobility skill for those jutting gold bronze rocks unlocked with story progress so you have to come back later. You can get totally obscene amounts of mana during exploration though so you wont feel limited on unlocking skill tree skills for long if thoroughly exploring and fighting.