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what that means is if you want to just leave your game running while you do other stuff and not focus on it, then you should start the grandmapocalypse (so wrinklers start spawning)
if you're an active player and like clicking golden cookies and stacking buffs, then i recommend ignoring it
so when you should start the grandmapocalypse is totally dependant on your playstyle
All in all, you will always want at least the first level of it, because wrinklers.