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The cell where the mutation might happen needs to be in contact with the required plants.
For example, assuming I want to mutate an Ichorpuff in a Level 3 Garden and Dirt:
Cr=Crumbspore; Ew=ElderWort; X=EmptyCell;
[X][Ew][X]
[X][Cr] [X]
[X][Ew][X]
Each empty cell touches an ElderWort AND a Crumbspore, fulfilling the pre-requisites and has a 0,2% chance to grow an Ichorpuff every time the garden ticks.
7%/tick while mature only.