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Red - A Special Team node
Blue - Squirrel Trainer node
Never seen one in Act 1 or Act 4. Do note that due to madness/corruption, if you rescue a pet during your run, you can in fact get the upgraded version by chance. So rescuing Betty in Act 1 can yield the upgrade immediately. Had a FANTASTICALLY easy run when I nabbed an upgraded Betty on Nez in Act 1.