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You plant a seed. Then the seed grows and in the end you can get a relic from it.
The fruit takes some real-world time to grow, but eventually will mature into celestial thaen fruit, which is plump and has yellow flesh. Harvest three of these, and the fourth time a celestial fruit would grow, the Ripened Heart relic grows instead. After this you can continue harvesting the fruits, but can't have more than one in your inventory. The tree is permanent for the character you grow it with, and will persist across rerolls.