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If you take another hit, the grey health from before is lost, essentially replaced by the most recent damage. For example, if you're hit for 70 damage, you'd have 35 points of grey health. Being hit by another attack for 20 damage afterwards would result in that 35 points of grey health being lost, and being replaced by 10 points of grey health.
The Backbone perk allows you to take 1-2 more hits without losing previous grey health. This means you can be struck multiple times, and end up healing the total grey health of the previous 3 hits instead of just 1.
Grey health is a common concept in video games.
So if an attack has an 80% grey health ratio, and you take a hit that does 100 damage, then you have lost 100 health. However, 80 points of that 100 damage is grey, meaning that under certain conditions, you can heal it back. There are usually other conditions that make it disappear completely too.
In Street Fighter, you could do a focus attack, and you would take damage as grey health. It would slowly heal itself over time, but if you took another hit, it would disappear entirely.
In Bloodborne, when you took a hit, you had some grey health left behind for a short time. If you kept attacking an enemy during that time, you could restore that grey health.
Bleeding in Monster Hunter works the same way, at least in MHW. You take damage over time, but when it is cured, that grey health will heal, or certain recovery item will instant heal all grey health. But if you take a hit before it regens that grey health disappears.
So if you have 200 max health, take a hit with 100 damage but that leaves 80 grey health then:
Your current Health = 100
the amount of health you can heal = 80
How much health you will have if you don't get hit again = 180
What back bone does is lets you take 2 hits instead of one, then three hits with the upgrade, before your grey health gets wiped out and is no longer able to heal itself.
Grey health is health that can regen by itself that you can get back.
grey health +2 is really strong imo, the description is not clear.
Its like this:
You get hit, you see your HP turns grey for a part of the damage you took.
You take a hit again, the grey dissapears along with more of your HP.
The passive gives essentially an extra hit while you already have grey health present. So like a fake extra small HP bar.
Grey health is health lost that can Regen. You seemed to understand this.
If you get hit again it disappears. Even if you had 100 grey health, and only got hit for 1 damage, all 100 points of your grey health would disappear on top of that 1 point of damage. This is the part that you seemed confused about.
The skill basically means that you won't lose that 100 grey health when you take the additional hit for 1 damage.
So without the skill:
50 hp + 100 grey health -> take 1 point of damage -> 49 hp + 0 grey health
With the skill:
50 hp + 100 grey health -> take 1 point of damage -> 49 hp + 100 grey health -> take 1 more point of damage -> 48 hp + 0 grey health